THE EARLY EOCENE FLORA OF HORSEFLY, BRITISH COLUMBIA, CANADA AND ITS PHYTOGEOGRAPHIC SIGNIFICANCE

About forty species, including a bryophyte, ferns, conifers, Ginkgo, and over 35 angiosperms, are recognized based on compression-impression remains from the early Eocene of Horsefly, British Colombia, Canada. This flora is in the north central part of a chain of late early Eocene fossil assemblages known as floras of the “Okanagan Highlands” (= “Okanogan High-lands” in the US). These floras extend from north central British Columbia, Canada southeast to Republic, Washington, USA. The Horsefly flora shows similarities to other Eocene Okanagan Highlands floras, such as McAbee, Falkland, Thomas Ranch and Republic, but with some additional rare taxa. In the broader sense, the Horsefly flora can be compared with early and middle Eocene floras of eastern Asia and midcontinental North America, but shares fewer elements with Europe. Ginkgo, Metasequoia, Pinus, Palaeocarpinus, ulmaceous leaves and Deviacer are shared with northeastern China as well as western North American sites. Fagopsis, Macginitiea, Dipteronia, Florissantia, Sassafras and Lagokarpos also occur in other western North American Eocene floras and Koelreuteria and Jenkinsella with Eocene paleofloras of China. These comparisons demonstrate probable floristic exchange between northwestern North America and northeastern China via a Beringian route during the late early Eocene.