Hunting, mating, patrolling, and rank trajectories for each male in the study, ordered by observed mating frequency, from most (A) to least (D). We pooled behavioral data into 5-year bins to visualize change over time while minimizing noise. Mating rates are scaled relative to the highest mating rate observed during that time interval. Hunting represents the proportion of hunts on which an individual participated. Patrolling represents the proportion of patrols for which the male was present. Rank represents the mean cardinalized Elo score. Here, we include PX, who suffered a scrotal wound at a young age and was effectively castrated. We excluded PX from analyses because he was not a possible sire.